1.Wuxi Fisheries College,Nanjing Agricultural University;2.National Demonstration Center for Experimental Fisheries Science Education,Shanghai Ocean University;3.Key Laboratory of Freshwater Fisheries and Germplasm Resources Utilization,Ministry of Agriculture and Rural Affairs,Freshwater Fisheries Research Center,Chinese Academy of Fishery Sciences,Wuxi

：National Freshwater Genetic Resource Center(FGRC18537)；Central Public-interest Scientific Institution Basal Research Fund, Freshwater Fisheries Research Center, CAFS (2023JBFM09, 2023TD65)