Annual changes of ovarian development and sex steroid levels in female cultured largefin longbarbel catfish (Hemibagrus macropterus) broodstocksLI Zuyang1,3,#, LI Shuang2,#,YUE Huamei1,YE Huan1,CHEN Yuankun2,RUAN Rui1,XIE Yong2,*,LI Chuangju1,*