National Key Research and Development Program of China (2022YFD2400502; 2022YFD2400103); National Modern Agricultural Industry Technology System (CARS-47-G31) from the Ministry of Finance and Ministry of Agriculture and Rural Affairs; Funding projects:Shandong Provincial Key Research and Development Program (2022LZGC016); The Key Research and Development Project of Hainan Province (ZDYF2025SXLH002); Qingdao Science and Technology Benefiting the People Demonstration Project (24-1-8-xdny-3-nsh);Central Public-Interest Scientific Institution Basal Research Fund (20603022024013); Chinese Academy of Fishery Sciences Research Operating Funds (2020TD19)