1.Huzhou University School of Life Sciences,Huzhou.Yangtze River Fisheries Research Institute,Wuhan, Hubei;2.Yangtze River Fisheries Research Institute,Chinese Academy of Fisheries Sciences;3.Huzhou University School of Life Sciences,Huzhou

National Key R&D Program Project "Impact Mechanism and Comprehensive Suitability Evaluation of Physical Habitat Changes on Fish in the Upper Yangtze River" (2023YFC3205903); Special Fund for Basic Scientific Research Funds of Central Public Welfare Research Institutes of the Chinese Academy of Fisheries Sciences (2023TD09); Scientific Research Project Funding of China Three Gorges Construction Engineering (Group) Co., Ltd. (Contract No.: JG/18056B, JG/18057B)