1. Key Laboratory of Freshwater Aquatic Genetic Resources, Ministry of Agriculture and Rural Affairs, Shanghai Ocean University, Shanghai 201306, China;2. Centre for Research on Environmental Ecology and Fish Nutrion of the Ministry of Agriculture and Rural Affairs, Shanghai Ocean University, Shanghai 201306, China;3.College of Marine and Bioengineering, Yancheng Institute of Technology, Yancheng Jiangsu 224000, China

The National Natural Science Foundation of China (No. 32273154), The Natural Science Foundation of Shanghai (22ZR1427300), The earmarked fund for CARS-48