1.Shanghai Ocean University,College of Marine Living Resources and Management;2.Yellow Sea Fisheries Research Institute,Chinese Academy of Fishery Sciences;3.Shandong Modern Fisheries Corporation

Natural Science Foundation of Shandong Province, China (No. ZR2022MD086), the Special Financial Projects of the Ministry of Agriculture and Rural Affairs for the Survey and Monitoring of Fish Spawning Grounds in the Bohai Sea and Yellow Sea (No. C1250515); National Key Basic Research Program of China (2015CB453302);Taishan Industrial Experts Programme (No. tsls20231201); Innovation Team of Fishery Resources and Ecology in the Yellow Sea and Bohai Sea (2020TD01)